Evolutionary studies are usually based on dichotomously branching phylogenetic trees. Russell et al. (pp. 37–56) use DNA data from several plastid and low-copy nuclear genes to generate phylogenetic networks of Polystachya, showing reticulate evolution. One group of allotetraploid species has a recent history of long-distance dispersal and multiple hybrid origins; another shows considerable morphological diversity on Madagascar and the Comoros.
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