The genus Cleome (Cleomaceae) includes more than 200 species including one, C. gynandra, that is known to be C4 and two others that are believed to be. Koteyeva et al. show that all three have features of C4 photosynthesis in their leaves and cotyledons. They are NAD-malic enzyme-type C4 species, and two very different forms of leaf morphology and Kranz anatomy have evolved in the genus, with adaptations to different environments that are supported by biogeographical information.
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