Eragrostis walteri (Poaceae), a grass with C3 foliar anatomy in an otherwise C4 genus and subfamily, has long been thought to be an exceptional example of the rare evolutionary reversion from C4 to C3 photosynthesis. Ingram et al. refute this hypothesis through phylogenetic analyses of DNA sequence data, which demonstrate that E. walteri has been misclassified and in fact belongs in the C3 Arundinoideae.