Annals of Botany Evolutionary history of sea barleys (Hordeum marinum) Hordeum marinum (sea barley) is a complex species that is usually recognized as having two subspecies, marinum (2n = 2x = 14) and gussoneanum (2n = 2x =14 and 2n = 2x = 28). Carmona et al. use comparative physical... botanyoneMay 9, 2014